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Re: Pine hybrids and pollen?



I will try to send a file containing a list of references on pine hybrids
that I compiled for an in house paper. If you don't get it and are interested
just write to me at P.O. Box 216, Rincon, Georgia, 31326 USA.
Best wishes

Marvin Zoerb
 


A  BIBLIOGRAPHY OF PINE HYBRIDS

Compiled by M. H. Zoerb, Jr.
10 April 1995


Adams, W.T., and Coutino, S., 1977. Isoenzyme markers useful for studies of the Pinus rigida x P.
taeda hybrid. Proc. 24th Northeast. For. Tree. Imp.  Conf. College Park, Maryland. pp. 1-13.

Ahn, K.Y., 1963. Studies on interspecific hybridization in the subgenus Diploxylon of genus Pinus.
Inst. For. Gen. Res. Rep. 3:29-43. Suwon, Korea.

Anderson, E., 1948. Hybridization of the Habitat. Evolution 2.1-9.

Anderson, E., 1953. Introgressive Hybridization. Biol. Rev. 28:280-307.

Anderson, E., 1968. Introgressive Hybridization. Hafner, Co., New York.

Anonymous, 1932.  Institute of Forest Genetics, Placerville, Ca. J.  of For. 30(5):623-624.
                                                                                                                                                       
Anonymous, 1990. Potential of slash pine hybrids for the lower coastal plain. Coop. For. Gen. Res.
Prog. , Univ. of  Fla., Gainesville, Fl.                                             .                                                
                    
Austin, L., 1927. A new enterprise in forest tree breeding. Jour. of Forestry 25:928-953.

Bailey, D.B., 1982. Shoot elongation in Pinus rigida x taeda hybrids. Silv.  Gen. 31(5-6)209-212.

Bannister, M.H., 1958. Hybridization between Pinus attenuata and P. radiata  in New Zealand.
Trans. of Royal Society of New Zealand. Vol.85, part 2.

Bannister, M.H., Brewerton, H.V., and McDonald, I.R.C., 1959. Vapour phase chromatography in
a study of hybridism in Pinus. Svensk Papperstidning 62(16) pp.567-573.

Barbour, H.F., 1975. Pond x loblolly hybrid test. Proc. 13th South. For.  Tree. Imp. Conf. Raleigh,
N.C. pp.105-112.

Barnes, B.V., and Bingham, R.T., 1962. Intermountain Forest and Range Expt.  Station. No. 104,
Juvenile performance of hybrids between western and eastern pines. 7 pp.

Barnes, R.D., and Mullin, L.J., 1978. Three year height performance of Pinus elliottii x P. taeda
hybrid families on 3 sites in Rhodesia. Silv. Gen.  27(6):217-223.

Bey, C.F., and Lorenz, R.W., 1970. Pitch x loblolly hybrid pines - mixed results from 7-year old
midwestern plantations. Univ. Ill. Agric. Stn. For. Res. Rep. 70-3, 4p.

Bingham, R.T., Squillace, A.E., and Patton, R.F., 1956. Vigor, disease resistance, and field
performance in juvenile progenies of the hybrid Pinus monticola x Pinus strobus. Forsgen. und
Forstplanz. now Sil. Gen.  5(4):104-112.

Bilan, M.V., 1953. Natural hybridization between loblolly and shortleaf pine in Texas. IUFRO Sec
22. Mtg. Zagreb, Yugoslavia pp. 123-125.

Blada, I. , 1992. Analysis of Genetic Variation in a Pinus strobus x  P. griffithii  F1 Hybrid
Population. Sil. Gen. 41:282-289.

Buchholz, J.T., 1944. The Cause of Sterility in Cross-Pollinations Between Species of Pines. Amer.
Jour. of Bot. 31.supl. 2pp.

Buchholz, J.T., 1945. Embryological aspects of hybrid vigor in pines.  Science 102(2641):135-142

Buchert, G.P., 1978. Breeding of other hard pines. Mimeo. p.137-144?

Buchert, G.P., 1983. Winter hardiness assessment in seedlings of pure and hybrid pines in south
Ontario. 28th N.E. For. Tree Imp. Conf. p.99-101.

Campbell, _, Hamaker, J.M., and Schmitt, D.M., 1969. Longleaf x shortleaf pine- a new hybrid.
Torrey Bot. Club Bull. 96:519-524.

Chapman,  H.H., 1922. A new hybrid pine. Jour. For. 20(7):729-734.

Callaham, R.Z., 1962. Resistance of three interspecific hybrids to blister rust. Jour. For.
60(6):409-410.

Callaham, R.Z., 1967. Hybridizing Pines With Diluted Pollen. Sil. Gen.  16:121-122.

Chiba, S., 1968. Heterosis in forest Tree Breeding. 9th sym.. Jap. Soc. of Breed. Tech. No. 69.

Conkle, M.T., and Tobolski, J.J., 1981.  Allozymes of linked loci segregate normally in seeds of an
Austrian x Japanese pine hybrid. Proc. 2nd North Central Tree. Imp. Conf. Lincoln, Ne. PP. 195-200.

Conkle, M.T., 1969. Hybrid population development and statistics. Sil. Gen.  18(5-6):197.

Cotton, M.H., 1972. Frequency of natural hybridization of shortleaf and loblolly pines in East Texas
as determined by morphological characteristics. Master's thesis. Sch. of  For. S. F. Austin Univ. 44p.

Critchfield, W.B., 1962. Hybridization of the southern pines in California.  For. Gen. Workshop Proc.
South. For Tree Imp. Comm. sponsored Publ.  22:40-48.

Critchfield, W.B., 1963. The Austrian X Red Pine Hybrid. Sil. Gen.  12:187-192.

Critchfield, W.B., 1965. Crossability and relationships of the big cone pines. USDA For. Ser. N.
Central For. Expt. Sta. Res. Paper No. 6.

Critchfield, W.B., 1965. A new hybrid Christmas tree. Southern Lumberman.  Dec.1965. p.132-133.

Critchfield, W.B., and E.L. Little, Jr., 1966. Geographic Distribution of the Pines of the World.
USDA Misc. Publ. 991.

Critchfield, W.B., 1967. Crossability Relationships of the Closed Cone Pines. Sil. Gen. 16:89-97.

Critchfield, W.B., and Krugman, S.L., 1967. Crossing the western pines at Placerville, California.
Univ. Wash. Arbor. Bull. 30(4):78-81,92.

Critchfield, W.B., 1975. Interspecific Hybridization in Pinus.:A Summary Review. Proc. 14th Mtg.
Can. Tree Imp. Assoc., pp. 99-105.

Critchfield, W.B., and B.B. Kinloch, 1986. Sugar Pine and It's Hybrids. Sil. Gen. 35:138-145.

Davidson, W.H., 1987. Pitch x loblolly hybrid pine performance on a West Virginia mine soil. Proc.
30th N.E. Tree Imp. Conf. p.96-101.

De la Punett, J.M., 1960. Hybridization and introgression between Pinus radiata and P. attenuata.
5th World Forestry Congress. Seattle, Wash.

Delmastro, R.J., 1976. Hybridization between several species of southern pines, their characteristics
and field performance. M.S. Thesis. N. C. State Univ.

Denison, N.P., Kietzka, J.E., . IUFRO Conf., Columbia, p.348-358.

Derr, H.J., 1966. Longleaf x slash hybrids at age 7 :survival, growth, and disease susceptibility. Jour.
For. 64:236-239.

Dorman, K.W., 1976. The genetics and breeding of southern pines. U.S. Forest Svc. Agric. Handb.
No. 471, Chap.15 pp. 295-325.

Duffield, J.W., and Righter, F.I., 1953. Annotated list of pine hybrids made at the Institute of Forest
Genetics. U.S. For. Svc. Cal. For. and Range Exp. Stn. Res. Note 86, 9p.

Duffield, J.W., 1952. Relationships and species hybridization in the genus Pinus. A. Forstgen.
Forstplanz. 1(4):93-99.

Duffield, J.W., 1954. The importance of species hybridization and polyploidy in forest tree
improvement. J. of For. 52(9):645-646.

Duffield, J.W., 1958. Benefits of hybridizing American forest tree species.
Jour. For. 56(11):809-815.

Duffield, J.W., and Snyder, F.B., 1958. Benefits From Hybridizing American Forest Tree Species.
J. For. 56:809-815.

Feret, P.P., Mayes, R.A., Tworkoski, Th. J., and Krebs, R.E., 1980.  Chromatographic analysis of
Pinus rigida x taeda hybrids and their parents. Silv. Gen. 29(3);155-156.

Florence, L.Z., 1973. Prospects for using natural loblolly x shortleaf hybrids for resistance to fusiform
rust. Proc. 12th South. For. Tree Imp. Conf. pp. 302-309.

Florence, L.Z., 1973. Predicting resistance to fusiform rust of loblolly, shortleaf, and natural loblolly
x shortleaf hybrid pines in East Texas.  Master's thesis. S. F. Austin Univ.

Fowler, D.P., 1978. Population improvement and hybridization. Unasylva 30(119-120):21-26.

Forukoski, T., and Sasaki,  M., 1982. Hybridization among species belonging to sub-section
Sylvestris and their resistance to wood nematode, including future cross breeding strategy. Kanto For.
Tree Breeding Inst. an. rpt.  No. 16. p.195-225.

Garrett, P.W., 1979. Hybridization in the genus Pinus. USDA For. Ser. Res.  Paper NE-436. pp.17.

Garrett, P.W., Kong, S.H., Shim, S.Y., Sohn, S.I., and Han, S.O., 1988. Growth of Korean hybrids
in test planting of  pitch x loblolly pine in the northeast United States. Res. Rpt. Inst. For. Gen. Korea
24:13-19.

Genys, J.B., 1971. Hybrids of Virginia pine x sand pine in Maryland.  Chesapeake Science
12:188-191.

Griffin, J.R., and Conkle, M.J., 1967. Early performance of knobcone and Monterey pine hybrids on
marginal timber sites. U.S. For. Svc. Res. Note.  PSW-156. 10p.

Grigsby, H.C., 1959. Two promising pine hybrids for the mid-south. South. Lumberman
198(2466):32-33.

Gyorfi, B., 1960. Hybrid vigor in forest trees and the genetic explanation of heterosis. Edeszeti Kut.
56(1/3):327-340.

Hall, R.C., 1959. Field tests of the resistance of hybrid pines to the pine reproduction weevil.  U.S.
Forest Service. Pacific Southwest Forest and Range Expt. Sta. Tech. Paper 33.

Haller, J.R., 1957. Taxonomy, hybridization and evolution in Pinus ponderosa and P. jeffreyi. Ph.D.
Thesis. Univ. of Cal. L.A. 169p.

Hamaker, J.M., and Schmitt, D.M.,1969. Longleaf  x shortleaf pine-a new hybrid. Bulletin of the
Torrey Botanical Club. Vol.96, No.5p. 519-524.

Hare, R.C., and Switzer, G.L., 1969. Introgression with shortleaf pine may explain rust resistance in
western loblolly pine. U.S. For. Svc. Res Note SO-88, 2p.

Harrington, T.A., 1953. Hybrid pines fail to grow as well as shortleaf, loblolly. Miss. Farm. Res.
16(4):8.

Haines, R.J., 1992. Mass propagation by cuttings, biotechnologies, and the capture of genetic gain.
IUFRO/AFOCEL Proc. Mass Prod. Tech. for Gen. Imp.  Fast Growing Species. pp.137-150.

Haines, R.J., and Walker, S.M., 1992. Mass vegetative propagation of slash x Caribbean
pine:operational status and research directions. IUFRO Conf.  p.488.

Haines, R.J., and Walker, S.M., 1993. Vegetative propagation for the capture of genetic gain with
hybrids in Queensland. Proc. Research Working Group 1 of the Australian Forestry  Council,
Canberra, ACT. Feb.1993.

Harding, K.J., 1994. Operational slash x caribea program in Queensland. Unpublished paper
presented as part of preliminary examination. N.C. State Univ.

Henry, B.W., and Bercaw, T.W., 1956. Shortleaf-loblolly hybrids free of  fusiform rust after five
years exposure. Jour. For. 54:779.

Hicks, R.R., 1974. Comparisons of seedlings from east Texas loblolly, shortleaf, and suspected
hybrids. Texas For. Ser. Paper No.24, six p.

Herrick, O.W., 1981. Economics of pitch x loblolly pine hybrids. Silv. Gen.  30(1):1-6.

Howcraft, N.H., 1974. A racial hybrid of Pinus merkusii, Trop. For. Res. Note SR28, Papua, New
Guinea.

Honeycutt, M., and Askew, G.A., 1989. Electrophoretic Identification of Loblolly Pine Shortleaf Pine
Hybrids. Sil. Gen. 38:95-96.

Hyun, S.K., 1958.  Feasibility of large scale pollination as a means of mass producing hybrid pine
seed. 10th Int.Cong. Gen. Proc. Vol. II:130-131.                                                                            
        
Hyun, S.K., 1959. Mass production of pitch-loblolly pine hybrid seed. Inst.  For. Gen. Suwon, 
Korea. Res. Ep. 1:11-24.

Hyun, S.K., and Ahn, K.Y., 1959. Principal characteristics of X Pinus rigitaeda. Inst. For. Gen.
Suwon, Korea. Res. Rep. 1:35-50.

Hyun, S.K., 1962. Improvement of pines through hybridization. IUFRO Proc.  13, Vol. 1(2),2p.

Hyun, S.K., 1969. The growth performance of pitch-loblolly hybrid pine produced by different
geographic races of loblolly pine in their early age. 2nd World Cons. on For Tree Brdg. FAO and
IUFRO vol. 1 pp.803-814.

Hyun, S.K., Ahn, A.K., and Hong, S.H., 1972. Developing Advanced Generation Breeding
Population for A Hybrid Breeding Program. Inst. For. Gen.  Suwon, Korea. Res. Pap. No.9, pp 1-8.

Hyun, S.K., 1974. The possibility of F2 utilization of Pinus rigida x taeda.  Korean  Jour. of
Breeding. 6:123-133.

Hyun, S.K., 1976. Interspecific hybridization in pines with special reference to P. rigidaxtaeda. Sil.
Gen. 25(5-6):188-191.

Jewell, F.F., 1961. Artificial testing of intra and interspecies southern pine hybrids for rust resistance.
Proc. 6th South. For. Tree. Imp. Conf.  pp.105-109.

Jewell,  F.F., 1961. Infection of artificially inoculated shortleaf pine hybrids with fusiform rust. Plant
Disease Reporter 45:639-640.

Jewell, F.F., 1985. Anatomical reaction to fusiform rust of interspecific progeny from crosses
between shortleaf and slash pine. Proc. IUFRO Conf. Rusts of hard pines. S2.06-10.U.S. For. Svc.
S.E.  For. Exp.  Stn.pp207-215.

Jinks, J.L., and Jones, R.M., 1958. Estimation of the components of heterosis.  Genetics
43(2):223-234.

Johnson, L.P.V., and  Heimburger, C., 1946. Preliminary Report on Interspecific Hybridization in
Forest Trees. Can. J. Res. 24:308-312.

Johnson, A.G., 1952. Spontaneous White Pine Hybrids. J. Arnold Arboretum. 33:179-185.

Johnson, A.G., 1955. Southern Pine Hybrids, Natural and Artificial. 3rd South. Conf. For. Tree Imp.,
New Orleans, La. pp.63-67.

Joly, R.J., and Adams, W.T., 1983. Allozyme analysis of pitch x loblolly pine hybrids produced by
supplemental mass pollination. For. Sci.  29:423-432.

Karfalt, R.P., Gerhold, H.D., and Palpant, E.H., 1975. The Possibilities for Interracial Hybridization
in Scotch Pine:Geographic Flowering Patterns and Crossability. Sil. Gen. 24.

Kleinschmit, J., Khurana, D.K., Gerhold, H.D., and Libby, W.J., 1993. Past, present, and anticipated
applications of clonal forestry. Clonal Forestry II, Springer-Verlag, Berlin. p.9-41.

Klotzsch, _ _, 1854. Abstract. The practical application of plant hybrids and cross breeds. Monthly
Newsletter of K. Academy of Science. Berlin pp.  535-562.

Knesick, D.R., Kuser, J.E., and Garrett, P.W., 1985. Supplemental mass pollination for the
production of southern pine hybrids. Proc. South. For.  Tree. Imp.Conf. Long Beach, Miss.
pp.187-193.

Krause, J.F., 1963. The Olustee arboretum, performance of 67 species of forest trees. U.S. For. Svc.
Stn. Paper SE-4.

Krause, J.F., Powers, H.R., Snow, G., 1982. Infection of shortleaf x loblolly pine hybrids inoculated
with C. quercum f. sp. echinatae and C. quercum f. sp. fusiforme. Phytopath. 72(4):431-433.

Krause, J.F., 1986. Breeding shortleaf x loblolly pine hybrids for the development of fusiform rust
resistant loblolly pine. South. Jour. of Apl.  For. 10:195-197.

Kriebel, H.B., 1972. Embryo development and hybridity barriers in the white pines(section Strobus).
Sil. Gen. 21(1-2):39-44.

Kuser, J.E., Knezick, D.R., and Garrett, P.W., 1987, Pitch x loblolly hybrids after 10 years in New
Jersey. N. Jour. of Applied For. 4(4):207-209.

Kang, K.W., 1966. Relationship between loblolly and pond pines in North Carolina. Ph.D. Thesis.
N.C. State. Univ. 135p.

Keng, H., and Little, E.F., 1961. Needle characteristics of hybrid pines.  Silv. Gen. 10:131-146.

Kim, C.K., 1963. The karyotype analysis in Pinus rigida, Pinus taeda, and their  F1 hybrid. Inst. For.
Gen. Res. Rep. 3:21-28. Suwon, Korea.

Kriebel, HB., 1963. Verifying species hybrids in the white pines. Proc.  For. Gen. Workshop, Macon,
Ga. Oct.25-27, 1962. SFTIC Publ. 22, pp.49-54.

Krugman, L., 1970. Incompatibility and inviability systems among some western North American
pines. Proc. Sex. Repro. Forest Trees, IUFRO, Sect.22, Finland, Part 2.

LaFarge, T., and Kraus, J.F. , 1977. Third year results of a shortleaf x loblolly hybrid progeny test
in Georgia. Proc. 14th South. For. Tree. Imp.  conf. pp.63-69.

LaFarge, T, and Hunt, D.L., 1980. Direction of pollination affects seed productivity in (shortleaf x
loblolly)x loblolly hybrids. USDA For. Ser.  S.E. Expt. Sta. Res. Note No. SE-294, 4pp.

LaFarge, T., and Kraus, J.F., 1980. A progeny test of (shortleaf x loblolly) x loblolly hybrids to
produce rapid growing hybrids resistant to fusiform rust. Sil. Gen. 29(5-6):197-200.                      

Lannor, R.M., 1974. Natural hybridization between Pinus edulis and P. monophylla in the American
southwest. Silv. Gen. 23(4):106-116.

Lester, D.T., 1973. The role of interspecific hybrids in forest tree breeding. Proc. 14th Mtg. Can.
Tree Imp. Assoc. pp.85-97.

Libby, W.J., 1958. The backcross hybrid Jeffrey x (Jeffrey x Coulter) pine.  Jour. For.
56(11):840-842.

Little, E.,  and Critchfield, . W.B., 1969. Subdivisions of the genus Pinus.  U.S. Forest Service Mis.
Pub. 1144.

Little, E.L., 1960. Designating hybrid forest trees. Taxon 9:225-231.

Little, E.L., Little, S., and Doolittle, W.T., 1967. Natural hybrids among pond, loblolly, and pitch
pines. U.S. Forest Svc. Res. Paper. NE-67 22p.

Little, E.L.,  and Richter, F.I., 1965. Botanical description of forty artificial pine hybrids. Tech. Bull.
No. 1345.

Little, S., and Somes, H.A., 1951. No exceptional vigor found in hybrid pines tested. Northeast For.
Expt. Sta. Res. Note No. 10.

Little, S., and Trew, I.F., . Progress report on testing Pitch x loblolly pine hybrids and on providing
hybrid seed for mass planting. Proc. 24th Northeast. Tree Imp. Conf. College Park, Maryland. pp.
14-28.

Little, S.,And Trew, I.F., 1976. Breeding and testing pitch x loblolly pine hybrids for the Northeast.
Proc. 23rd Northeast For. Tree Imp. Conf., pp.71-85.




Little, S., and Trew, I.F., 1979. Pitch x loblolly hybrids: loblollies for the north. Jour. of For.
77:709-716.

Long, E.M., 1973. Performance of interspecific hybrids and exotic pines in Texas. Proc. 12th South.
For. Tree. Imp. Conf. pp.293-301.

Lorenz, R.W., and Spathe, J.N., 1953. The growth of two hybrid pines tested in southern Illinois.
Univ. Ill. Agric. Exp. Stn. For. Note 38 2p.

Lotan, J.E., 1967. Early performance of Pinus contorta x banksiana hybrids.  Forestry Abstracts,
Vol.19(1):70.

Martin, B., 1988. The benefits of hybridization. How do you breed for it?.  AFOCEL-UAIC pp.20.

McWilliams, J.R., 1958. Pollination, pollen germination and interspecific hybridization in Pinus. Ph.D.
thesis, Yale Univ.

McWilliams, J.R., 1959. Interspecific Incompatibility in Pinus.  Amer. Jour.
Bot. 46:425-433.

Mergen, F., 1954. Tree Improvement Research at the Lake City, Florida, Research Center, A project
analysis. U.S. For. Svc. Stn Paper. No. 45.

Mergen, F., 1957. Genetic variation in needle characteristics of slash pine and some of its hybrids.
Silv. Gen.7(1):1-40.

Mergen, F., 1958. Genetic variation in needle characteristics of slash pine and some of its hybrids.
Sil. Gen. 7:1-9.

Mergen, F., Stairs, F.W., and Snyder, E.B., 1965. Natural and controlled loblolly x shortleaf hybrids
in Mississippi. For. Sci. 11:306-314.

Mergen, F., 1959. Applicability of the distribution of stomata to verify pine hybrids. Sil. Gen.
8(4):107-109.

Mergen, F., and Furnival, G.M., 1960. Discriminate Analysis of Pinus thunbergii x P. densiflora
hybrids. Proc. Soc. Am. For. p.36-40.

Meyer, H., 1954. Der Bastard Pinus strobus x excelsa. IUFRO Proc. 1953, Sec.22, No.13, Rome,
Italy. 2 pp.

Miller, J.M., 1950. Resistance of pine hybrids to the pine reproduction weevil. For. Res. Note No.
68, Calif. For. and Range Expt. Sta.

Minckler, L.S., 1952. Loblolly pine seed source and hybrid tests in southern Illinois. U.S. For. Svc.
Cent. States For. Exp. Stn. Tech. Paper 128, 8p.

Morris, R.W., and Critchfield, W.B., 1980. The putative Austrian x red pine hybrid: A test of
paternity based on allelic variation at enzyme specifying loci. Silv. Gen. 29(3):93-99.

Moss, E.H., 1949. Natural  pine hybrids in Alberta. Can. Jour. Res. Sect. C.  Bot. Sci. 27:218-229.
species. Ph.D. thesis. Sch. of For. N.C. State Univ.

Namkoong, G., 1963. Comparative analyses of introgression in two pine species. Ph.D. thesis. Sch.
of For. N.C. State Univ.

Nelson, C.D., 1991. Fusiform rust incidence and volume growth in a first generation backcross
population, (shortleaf x slash)x slash. Proc. 21st South. For. Tree Imp. Conf. Knoxville, TN.
pp.152-159.

Nikles, D.G., 1964. Cross progenies of P. elliottii x P. caribaea. Austr.  For. Res. 1:32-33.

Nikles, D.G., 1966. Comparative variability and relationships of Caribbean pine and slash pine. Ph.D.
Thesis. N.C. State. Univ. 201p.

Nikles, D.G., 1981. Some successful hybrid breeds of forest trees and the need for further
development in Australia. Proc. 7th Meet. RWG No.  1--Forest Genetics, Traragon, Victoria

Nikles, D.G., and Robinson, M.J., 1989. The development of Pinus hybrids for operational use in
Queensland. In breeding tropical trees;population structure and genetic improvement strategies in
clonal and seedling forestry. Winrock Internet. USA and OFJ,UK.P272-282.

Nikles, D.G., Strategy and rationale of the breeding programmes with Pinus caribaea and its hybrids
in Australia. Proc. IUFRO Joint meeting of working parties on Breeding theory, progeny testing, and
orchards.  Williamsburg, Va. N.C. State Univ. pp.298.

Nikles, D.G., 1993. Hybrids of forest trees:the basis of hybrid superiority and a discussion of breeding
methods. Queensland Forest Service.

Nikles, D.G., Power, M.B., and Toon, P.C., 1993. Appropriate breeding methods to support clonal
forestry with P. elliottii x P. caribea var. hondurensis hybrids in Queensland. Clonal Forestry
Workshop. Queensland Res. Inst.  paper No.4, 21pp.

Panetsus, C.P., 1975. Natural hybridization between P. halepensis and P. brutia in Greece. Silv. Gen.
24(5-6):163-168.

Porterfield, J.D., 1991. The verification, description and inheritance patterns of putative P. virginiana
x P. clausa and P. clausa x P.  virginiana hybrid pines. Proc. 21st. South. For. Tree Imp. Conf. 
Knoxville, TN. pp.160.abstract.

Powers, H., Jr., and H.J. Duncan, 1976. Increasing Fusiform rust Resistance by Intraspecific
Hybridization. For Sci. 22(3):267-268.

Righter, F.I. and Stockwell, P., 1949. The Fertile Species Hybrid, Pinus murraybanksiana. Madrono
10:65-69.

Righter, F.I., and Duffield, J.W., 1951. Hybrids between ponderosa and apache pine. J. of For.
Vol.49, no. 5. p345-349.

Righter, F.I., and  Duffield, J.W., 1951. Interspecies Hybrids in Pines, A Summary of Interspecific
Crossing in the Genus Made at the Institute of Forest Genetics. J. Hered. 42:75-80.

Righter,  F.I., 1955. Possibilities and limitations of  hybridization in Pinus. Proc. 3rd South. Conf.
For. Tree. Imp., New Orleans, La. pp54-63.

Righter, F.I., 1961? . Forest tree improvement through inbreeding and intraspecific and interspecific
hybridization. Proc. 5th World Forestry Cong.

Righter, F.I., 1962. Evidence for hybrid vigor in forest trees. Tree Growth.  pp. 345-355. Ronald
Press.

Sax, K., 1960. Meiosis in Interspecific Pine Hybrids. For. Sci.  6(2):135-138.

Saylor, L.C.,  and Smith, B.W., 1966. Meiotic irregularity in species and interspecific hybrids of
Pinus. Am. J. Bot. 53:453-468.                                                                                                       
    
Saylor, L.C., 1967. Variability of chromosome structure and behavior in southern pine hybrids. 9th
South. For. Tree Imp. Conf. Proc. pp. 95-100.

Saylor, L.C., 1972. Karyotype analysis of the genus Pinus-subgenus Pinus.  Sil. Gen. 21:155-163.

Saylor, L.C., 1969. Chromosomal differentiation as a barrier to interspecific hybridization among
pines. 2nd World Consult. on For. Tree. Brdg.  Washington, D.C.  6p.

Saylor, L.C., and Zobel, B.J., 1973. Interspecific hybridization involving sand pine--past attempts and
future potential, p.244-253.In sand pine symposium proc. U.S. For. Svc. Tech. Rep. SE-2.

Saylor, L.C., and Kang, K.W., 1973. A study of sympatric populations of Pinus taeda and Pinus
serotina in North Carolina. Jour. Elisha Mitchell Sci. Soc.  89(142):101-110.

Saylor, L.C., and Koenig, R.L., 1967. The slash x sand pine hybrid. Sil. Gen.  16(4):134-138.

Schmidtling, R.C., and Scarborough, N.M., 1968. Graphic analysis of Erambert's hybrid. U.S. Forest
Service Research Note No. SO-80.

Schmidtling, R.C., 1989. Wide crosses in loblolly pine:Crossing Livingston Parish with Atlantic coast
and central Florida populations. Proc. 20th South. For. tree. Imp. Conf. Charleston, S.C. pp.309.

Schmitt, D.M., 1968. Performance of southern pine hybrids in southern Mississippi.  U.S. Forest Svc.
Res Paper SO-36 15 p.

Schmitt, D.M., 1969. Nanism in slash x shortleaf pine hybrids. For. Sci.  15:174-175.

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Schoenike, R.E., Van Lean, D.H., and Benson, J.D., 1977. Comparison of shortleaf, loblolly, and
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Slee, M.U., 1971. Natural Hybridization between Cuban Caribbean and Slash Pines. Sil. Gen.
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Stebbins, G.L., 1969. The significance of hybridization for plant taxonomy and evolution. Taxon
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Woessner, R.A., 1972  Crossing among loblolly pines indigenous to different areas as a means of
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Woessner, R.A., 1972 Growth patterns of one year old loblolly pine seed sources and
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Wright, J.W., 1964. Hybridization between species and races. Unasylva 18(2-3):73-74

Wright, J.W., Lemmien, W.A., and Canavera, D.S., 1969. Abundant natural hybridization between
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Zavarin, E., Critchfield, W.B., and Snajberk, K., 1969. Turpentine composition of x Pinus contorta
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Zobel, B.J., 1951. The natural hybrid between Coulter and Jeffrey pines.  Evolution 5(4):405-413.

Zobel, B.J., 1951. Oleoresin composition as a determinant of pine hybridity.  Bot. Gaz. (Chicago)
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Zobel, B.J., 1953. Are there natural loblolly-shortleaf pine hybrids. Jour.  For. 51(7):494-495.

Zobel, B.J., Campbell, T.E., Cech, F.C., and Goddard, R.E., 1956.  Survival and growth of  native
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Zsuffa, S., 1973. Variation in Rooting Pinus strobus and P. griffithii x P. strobus Trees. Sil. Gen.
22:119-120.

Zsuffa, L., 1975. Broad sense heritability values and possible genetic gains in clonal selection of 
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